+44Kontinente:EUAS
1. Lebendfotos
1.1. Falter
1.2. Ausgewachsene Raupe
1.3. Jüngere Raupenstadien
1.4. Fraßspuren und Befallsbild
1.5. Puppe
1.6. Ei
2. Diagnose
2.1. Männchen
2.2. Weibchen
2.3. Genitalien
2.3.1. Männchen
2.3.2. Weibchen
2.4. Erstbeschreibung
2.5. Beschreibung von John Curtis als Hadena cucubali
3. Biologie
3.1. Habitat
3.2. Raupennahrungspflanzen
3.3. Parasitoide
4. Weitere Informationen
4.1. Andere Kombinationen
- Noctua rivularis Fabricius, 1775 [Originalkombination]
- Hadena rivularis (Fabricius, 1775) [so in Karsholt & Razowski (1996)]
- Sideridis rivularis (Fabricius, 1775) [bis Nedumpally et al. (2025) übliche Kombination]
4.2. Synonyme
- Noctua cucubali Denis & Schiffermüller, 1775
- Phalaena (Noctua) rivosa Ström, 1783
- Hadena behenis Freyer, 1845
4.3. Unterarten
- Aneda rivularis pacifica (Hacker, 1996)
4.4. Taxonomie
Nedumpally et al. (2025: 18) kamen in ihrer umfangreichen genetischen Studie - unter Einbeziehung diverser Gene der Kern-DNA - zum Schluss, dass die Gattung "Sideridis" im gegenwärtigen Umfang alles andere als monophyletisch ist. Sie liefern keine komplette Neugliederung, sondern schlagen nur vor, "Sideridis rivularis" aus der Gattung herauszunehmen und (wieder) mit Aneda zu kombinieren, dessen Typusart sie ist. Es ist also zu erwarten, dass es hier noch zu weiteren Verschiebungen und Neukombinationen kommen wird. Die Autoren schrieben: "Third, the species composition of the genus Sideridis Hübner, 1821 as delimited, for example, in Hacker et al. (2002), Aarvik et al. (2017) and Top et al. (2023) clearly needs further study, as it was not found to be monophyletic in our analysis. In fact, Noctua rivularis Fabricius, 1775 (type species of Aneda Sukhareva, 1973) (Figure 6h) was recovered as sister to the clade comprising Conisania Hampson, 1905 and Hadena Schrank, 1802 with maximum statistical support (Figure 5), whereas Sideridis reticulata (Goeze, 1781) (Figure 6i) appeared to be sister to a far larger assemblage of taxa including Hecatera Guenée, 1852, Hada Billberg, 1820, N. rivularis, Conisania and Hadena, also with maximum statistical support (Figure 5). According to Hacker et al. (2002), S. reticulata is more closely related to S. lampra (Schawerda, 1913), the type species of Sideridis (originally fixed as Noctua evidens Hübner, 1808 a senior albeit unusable synonym of lampra, which is a junior primary homonym of a Thunberg’s name), and thus, we tentatively consider this part of the paraphyletic genus to be the ‘true’ Sideridis. As for N. rivularis, our data indicate that it has to be moved to another genus. In many earlier publications, this species has been associated with Hadena (e.g., Forster & Wohlfahrt, 1971; Nowacki & Fibiger, 1996; Skou, 1991), but our results confirm that such position is erroneous. Thus, we suggest reviving Aneda Sukhareva, 1973 from synonymy with Sideridis and treating it as a full genus (stat. rev.) according to Beck (1996) and Rákosy (1997) in order to avoid paraphyletic groups, resulting in a combination Aneda rivularis (Fabricius, 1775) (comb. rev.) for its type species."
(Autor: Erwin Rennwald)
4.5. Literatur
- Curtis, J. (1823-1840): British Entomology; Being Illustrations and Descriptions of the Genera of Insects Found in Great Britain and Ireland: Containing Coloured Figures from Nature of the Most Rare and Beautiful Species, and in Many Instances of the Plants upon which they are Found. Vol V. Lepidoptera, Part I. — [Not paginated]. London (E. Ellis & Co.).
- Ebert (1998) (= Ebert 7), 216-219.
- Erstbeschreibung: Fabricius, J. C. (1775): Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. 1-832 (Flensburg und Leipzig: Officina Libraria Kortii).
- Nedumpally, V., Zilli, A., Yapar, E., Tammaru, T., Lemmon, A.R. & E. Õunap (2025): Elaborating the phylogeny of Noctuidae by focusing on relationships between northern European taxa. — Systematic Entomology. 2025;e70010: 1-25. https://doi.org/10.1111/syen.70010.
